diff --git a/wp-includes/link-template.php b/wp-includes/link-template.php
index 2a1c7b2382..5a80a1cfde 100644
--- a/wp-includes/link-template.php
+++ b/wp-includes/link-template.php
@@ -2426,10 +2426,19 @@ function _navigation_markup( $links, $class = 'posts-navigation', $screen_reader
 	/**
 	 * Filter the navigation markup template.
 	 *
+	 * Note: The filtered template HTML must contain specifiers for the navigation
+	 * class (%1$s), the screen-reader-text value (%2$s), and placement of the
+	 * navigation links (%3$s):
+	 *
+	 *     <nav class="navigation %1$s" role="navigation">
+	 *         <h2 class="screen-reader-text">%2$s</h2>
+	 *         <div class="nav-links">%3$s</div>
+	 *     </nav>
+	 *
 	 * @since 4.4.0
 	 *
-	 * @param string $template The default template
-	 * @param string $class The class passed by the calling function.
+	 * @param string $template The default template.
+	 * @param string $class    The class passed by the calling function.
 	 * @return string Navigation template.
 	 */
 	$template = apply_filters( 'navigation_markup_template', $template, $class );
diff --git a/wp-includes/version.php b/wp-includes/version.php
index 25e0e69c2c..3282b1ff3e 100644
--- a/wp-includes/version.php
+++ b/wp-includes/version.php
@@ -4,7 +4,7 @@
  *
  * @global string $wp_version
  */
-$wp_version = '4.4-alpha-33716';
+$wp_version = '4.4-alpha-33717';
 
 /**
  * Holds the WordPress DB revision, increments when changes are made to the WordPress DB schema.